Diplopanax cacaoides

Diplopanax cacaoides

(Zenker) Manchester et M.E.Collinson

Fossil group Plants
Order Cornales
Family Mastixiaceae
Genus Diplopanax
Taxonomic status Accepted
Geological age Ypresian

Summary

Diplopanax cacaoides was an early Eocene mastixioid plant that produced a large, very thick-walled one-seeded fruit stone. The robust endocarp formed a substantial protective case around the seed, with a long germination valve and a curved internal seed cavity. No attached vegetative organs are known from these fruits, so the whole plant should not be reconstructed from them alone.

The London Clay material is especially important because it includes the oldest recognised record of Diplopanax. Some fossils were historically grouped under Mastixia cantiensis, but modern anatomical work showed that this old assemblage contained more than one kind of mastixioid fruit. The accepted identification therefore depends on the fruit’s internal construction rather than the old label.

Quick Identification

Look for: a large ovoid to cylindrical one-seeded mastixioid stone, typically 17–32 mm long, with an exceptionally thick wall and a horseshoe-shaped seed cavity; the long germination valve and internal wall anatomy are more reliable than an old identification as Mastixia cantiensis.

Diagnostic Features

  • Endocarp is unilocular and ovoid to cylindrical, rounded basally and rounded to truncate apically.
  • London Clay examples are about 17–32 mm long and 12–22 mm wide.
  • Locule is horseshoe-shaped in transverse section.
  • Endocarp wall is exceptionally thick, reaching about 9 mm.
  • Wall contains mainly radial fibres with scattered vascular bundles and secretory canals, without the median fibrous septum of Tectocarya grandis.

Fossil Details

Typical size
About 17–32 mm long and 12–22 mm wide.
Fossil material
Fruit
Preservation
Thick pyritised endocarps may break into incomplete sections, obscuring the original outline and valve. Historical material labelled Mastixia cantiensis is heterogeneous, so specimens should not be transferred to this species solely on an old name; internal anatomy and locule construction should agree.

Similar Species

Recorded Fossil Sites

No fossil-site occurrences have yet been recorded for this species.

Synonyms & Previous Names

Mastixia cantiensis

References

  • Steven R. Manchester; Margaret E. Collinson. (2022). Mastixioid fruits (Cornales) from the early Eocene London Clay Flora: morphology, anatomy and nomenclatural revision. Fossil Imprint, 78(1), 310–328 View reference
Last Reviewed
26 August 2026 Reviewed by Sam Harris