Squalus minor
Squalus minor
Common Name: Dogfish Shark
Order: Squaliformes
Family: Squalidae
Genus: Squalus
Geological Age: Ypresian
Summary
Squalus minor is a small extinct dogfish shark known from Palaeocene and Eocene marine deposits. Its tiny teeth are quite different from the familiar upright teeth of many sharks, being low, broad and strongly tilted towards one side.
The teeth have a smooth cutting edge, a rounded heel behind the main point and a characteristic V-shaped projection of the crown extending down across the front of the root. Complete teeth are usually only a few millimetres across and can therefore easily be missed when collecting or processing fine material.
Squalus minor occurs across a long interval from the Early Palaeocene to Middle Eocene and is known from the London Basin, including the London Clay Formation. Because different Squalus species have extremely similar teeth, complete specimens showing the crown, root and central projection provide the most reliable identifications.
Geological Age & Range Notes
Early Palaeocene to Middle Eocene — approximately Danian to Lutetian. Published records indicate a range from calcareous nannofossil zones NP3 to NP16.
Within the London Basin, Squalus minor occurs in both Palaeocene and Early Eocene deposits.
Fossil Material
Known Localities
Diagnostic Features
- Teeth are very small, generally only a few millimetres wide.
- The tooth is normally wider than it is tall, giving it a low, compact appearance.
- The main point is strongly tilted towards one side rather than pointing straight upwards. This sideways pointing shape is typical of dogfish teeth.
- The front edge of the main point is usually long and gently rounded or curved.
- Behind the main point is a low, rounded raised area or “heel” rather than a separate pointed side cusp.
- There is a small notch separating the main point from this rounded heel.
- The cutting edges are normally completely smooth, with no serrations.
- The crown surface itself is also generally smooth, without the strong lines, folds or decoration seen on some other fossil shark teeth.
- On the front face of a complete tooth there is a very noticeable V-shaped or tongue-shaped piece of enamel extending down over the root. This is called the apron, but to the eye it simply looks like the centre of the crown continues downwards below the rest of the tooth.
- This central projection can extend below the bottom of the root and is one of the most useful features when recognising Squalus teeth.
- On the opposite side of the tooth there is a much smaller central projection above a large opening in the root.
- The root is low and fairly flat, rather than being divided into the long root lobes seen in sharks such as Striatolamia.
- Complete side teeth can show small extensions of the root at either end.
- Teeth change relatively little around the mouth compared with many other sharks. Upper teeth tend to be slightly narrower than lower teeth, but the same basic sideways pointing shape is retained.
Identification Notes
Squalus minor is an extinct member of the dogfish shark family and its teeth have the characteristic low, sideways pointing appearance seen in Squalus.
At first glance they can look rather unlike a conventional shark tooth because the point does not rise vertically from the middle. Instead, the crown appears to have been pushed over towards one side.
A good Squalus minor tooth should therefore have:
a low broad crown + sideways pointing main point + rounded heel behind it + completely smooth edges + broad central projection extending down the front of the root.
Look at the central projection
One of the easiest features to check is the front of the tooth.
In the centre of the crown is a prominent V-shaped or rounded projection that extends down across the root.
On a complete S. minor tooth this can reach below the bottom edge of the root.
This feature is characteristic of Squalus teeth and is especially useful when dealing with very small specimens where the shape of the main point may be difficult to see.
Look at the rounded heel
Behind the main point is a high, smoothly rounded heel.
It should not look like a separate sharp side point. Instead, the cutting edge drops from the main point into a small notch and then rises slightly into a broad rounded area.
The combination of a slightly curved main point and high rounded heel was one of the features used by Leriche to characterise S. minor.
Check for serrations
The cutting edges of Squalus minor should normally be smooth.
This is useful when separating it from some similar fossil dogfish teeth.
Squalus minor vs Squalus orpiensis
These two Palaeogene dogfish sharks can occur in deposits of similar age and their teeth can look quite similar.
Squalus minor generally has:
- smooth cutting edges
- a comparatively compact crown
- a rounded rear heel
- a smooth base to the crown
Squalus orpiensis generally has:
- a longer and more stretched-out tooth
- cutting edges that can show fine serrations
- fine wrinkles or crenulations along the inner base of the crown
- larger openings in parts of the root
These differences are useful, particularly when complete teeth from the same deposit can be compared.
Squalus smithi
The name Squalus smithi appears in older literature.
Modern work considers S. smithi to be a younger synonym of Squalus minor because the main distinction originally used between them was tooth size, and size alone is not considered reliable enough to separate the two species.
Species identification needs some caution
There is an important limitation with Squalus minor.
The teeth of living species of Squalus can be extremely difficult to distinguish from one another, even when complete sets of teeth are available. Fossil species are normally known only from isolated teeth.
Researchers have therefore cautioned that fossils currently grouped under the name Squalus minor may not necessarily represent a single biological species.
For the identification database, a well preserved tooth from an appropriate Palaeocene or Eocene deposit showing the classic S. minor morphology can reasonably be identified as Squalus minor.
If the crown is badly worn, the root is missing, or the important features cannot be seen clearly, Squalus sp. or Squalus cf. minor would be safer.
Quick recognition
Look for:
tiny tooth + wider than tall + strongly sideways pointing cusp + rounded heel + smooth cutting edges + central V-shaped projection over the root.
References & Literature
- Leriche, M. (1902). Les poissons paléocènes de la Belgique. Includes the first valid description and illustrations of the species as Acanthias minor. The figured specimens form the important reference material for S. minor.
- Casier, E. (1966). Faune ichthyologique du London Clay. British Museum (Natural History). Major reference for the shark and fish fauna of the London Clay Formation.
- Herman, J. (1977). Les Sélaciens des terrains néocrétacés et paléocènes de Belgique et des contrées limitrophes. Important treatment of Palaeocene sharks including Squalus.
- Ward, D.J. (1980). The distribution of sharks, rays and chimaeroids in the English Palaeogene. Tertiary Research 3(1): 13–19. Important reference for British Palaeogene shark distribution.
- Cappetta, H. (2012). Chondrichthyes: Mesozoic and Cenozoic Elasmobranchii: Teeth. Handbook of Paleoichthyology, Volume 3E. Major modern reference for fossil shark tooth morphology and taxonomy.
- Migom, F., Christiaens, Y. & Deceuckeleire, M. (2021). The Lower Eocene (Ypresian) Chondrichthyes from Egem, Belgium. Part 1. Includes discussion of Squalus minor and the synonymy of S. smithi.
- Rodríguez, D., Ward, D.J. & Quezada, J.A. (2023). Paleontology and stratigraphic implications of a late Paleocene elasmobranch assemblage in Talcahuano, southcentral Chile. Andean Geology 50(2): 217–247. Contains a modern description and detailed discussion of Squalus minor tooth morphology and range.
- Pollerspöck, J., Straube, N. & Reinecke, T. (2025). Nomenclatural revision of the squaliform taxa introduced by Daimeries (1888). Paleoichthys 9: 1–7. Establishes Leriche (1902) as the authorship of Squalus minor under modern nomenclatural rules.